Cereal Biotechnology - Peter C. Morris , James H. Bryce.pdf

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Cereal biotechnology
Edited by
Peter C Morris and James H Bryce
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Published by Woodhead Publishing Limited
Abington Hall, Abington
Cambridge CB1 6AH
England
Published in North and South America by CRC Press LLC
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Boca Raton FL 33431
USA
First published 2000, Woodhead Publishing Limited and CRC Press LLC
2000, Woodhead Publishing Limited
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1
Introduction
P. C. Morris and J. H. Bryce, Heriot-Watt University, Edinburgh
1.1 Cereals: an introduction
Cereals owe their English name to the Roman goddess Ceres, the giver of grain,
indicative of the antiquity and importance of cereals (Hill 1937). This
importance is still very much the case today; cereals of one sort or another
sustain the bulk of mankind’s basic nutritional needs, both directly and
indirectly as animal feed. It is primarily the grains of cereals that are useful to us,
although the vegetative parts of the plant may be used as fodder or for silage
production, and straw is used for animal bedding.
Cereals are members of the large monocotyledonous grass family, the
Gramineae . The flowering organs are carried on a stem called the rachis, which
may be branched, and in turn bears spikelets which may carry more than one
flower at each node of the rachilla (Fig. 1.1). The spikelets may be organised in a
loose panicle as in sorghum, oats and some millets, or in a tight spike, as in
wheat. The length of the internodes of the rachis and of the rachilla, and the
number of flowers at each node of the spikelet determine the overall
architecture. Each spikelet is subtended by two bracts or leaf-like organs
termed the glumes, and each flower in the spikelet is enclosed in two bract-like
organs called the lemma and palea. The lemma may be extended to form a long
awn. In some cereals or cereal varieties the lemma and palea may remain
attached to the grain; these are termed hulled or husked grains, such as oats and
most barleys, as opposed to naked grains such as most wheats and maize (Fig.
1.1).
The cereals, with the exception of maize, are dioecious. Each flower bears
both male organs; the three anthers (six in rice), and female organs; the ovary
which carries two feathery stigmas. In maize, the male flowers are borne in
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